On Easter Island, a young rabbit named E.B. (Russell Brand) is intended to succeed his father Mr Bunny (Hugh Laurie) as the Easter Bunny. Intimidated by the calling's demands, E.B. runs away to Hollywood to pursue his dream of becoming a drummer. Mr Bunny sends his three ninja royal guards, the Pink Berets (Janet Healy), to find his son. Meanwhile, Easter Chick Carlos (Hank Azaria) and his music-obsessed assistant, Phil (also Hank Azaria), plot a coup d'état against Mr Bunny to take over the Easter organization.
Wandering through Los Angeles, E.B. is hit by Fred O'Hare (James Marsden), an out of work slacker who was driving to his sister Sam's (Kaley Cuoco) boss's house he's housesitting . Feigning injury, E.B. persuades Fred to take him in as he recovers, but when E.B. causes trouble Fred attempts to abandon him in the wilderness. E.B. persuades the human to help him by claiming to be the Easter Bunny, whom Fred saw in operation in his youth. The rabbit sees the Berets closing in on him and hides inside a business where Fred is having job interview. E.B. enjoys a successful recording session with the Blind Boys of Alabama as their substitute drummer, but ruins Fred's job interview. In the process, E.B. gets a tip about a possible audition for David Hasselhoff, who invites him to perform on his show.
Afterward, Fred attends his adopted younger sister Alex's (Tiffany Espensen) Easter school pageant with E.B. hiding in a satchel. E.B., alarmed that the Pink Berets have apparently found him and disgusted by Alex's awful rendition of Here Comes Peter Cottontail, dashes out and disrupts the show. Fred feigns a ventriloquist's act with the bunny's cooperation as his dummy and they lead the show in singing, "I Want Candy". Both his father, Henry (Gary Cole) and Alex, are angry about the upstaging, but Fred is inspired to be the Easter Bunny himself. E.B. is skeptical, but he agrees to train the human and finds that Fred has some genuine talent for it.
As the Pink Berets close in on him, E.B. prepares a decoy to fake his death and leaves for Hasselhoff's show. The Berets see the decoy and, horrified that the human has apparently killed E.B., capture Fred and take him to Easter Island. Fred is held captive and confronted by Mr Bunny and Carlos about killing E.B. Carlos pretends to be upset about E.B.'s death, ignores Fred's protests about the truth of the decoy and seizes control of the Easter factory.
Meanwhile, at the Hoff Knows Talent live show E.B. is in his dressing room preparing for his performance but his reflection begins to berate him for leaving Fred. Just then a production assistant (also Russell Brand) arrives to tell him that he is next to perform. E.B. leaves his dressing room and begins to feel guilty about leaving Fred so he discusses the situation with Hasselhoff, who advises him to go back and help his friend. E.B. finds evidence of Fred's capture and races back to the factory. He confronts Carlos, but is immobilized in gummi candy and tossed into the chocolate bunny carving line. Fred and Mr Bunny are tied up with black licorice and are to be boiled alive. Fred eats his way out and E.B. survives by dodging the blades. Carlos, now a chick-bunny combination due to Easter magic engages battle with E.B. and defeats him easily due to his size and then tries to lead the Easter Egg sleigh out with Phil directing via lighted wands.
To stop the launch, E.B. improvises a drum session. Uncontrollably driven to dance to the beat, Phil provides the wrong signals, causing a crash and Carlos is subdued. After being defeated, Carlos is made to pull the Easter Egg sleigh and Phil is made the new head of the Easter Chicks. E.B. and Fred are made co-Easter bunnies and take over the job. Fred's family is now proud that he has a job and responsibilities.
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